These articles have been published through the work of the All Out Africa Research Unit and are listed in date order:

African Savanna Publications:

Associations of avian facial flushing and skin colouration with agonistic interaction outcomes.

Bamford, A.J., Monadjem, A. & Hardy, I. 2010. Ethology 116: 1-8.


Detection rates of ungulates in the eastern Swaziland lowveld.

Collier, B., McCleery, R., Calhoun, K., Silvy, N. Roques, K.G, & Monadjem, A. 2011. South African Journal of Wildlife Research 41: 61-67.


Impact of crop cycle on movement patterns of pest rodent species between fields and houses in Africa.

Monadjem, A., Mahlaba, T.A., Dlamini, N., Eiseb, S.J., Belmain, S.R., Mulungu, L.S., Massawe, A.W., Makundi, R.H. & Taylor, P.J. 2011. Wildlife Research


Long-term changes in vegetation and bird communities in southern African savannas.

Serami, C. & Monadjem, A. 2011. Diversity & Distributions 1-11. doi: 10.1111/j.1472-4642.2011.00810.x


Molecular diet analysis of two African free-tailed bats (Molossidae) using high throughput sequencing.

Bohmann, K., Monadjem, A., Noer, C.L., Rasmussen, M., Zeale, M.R.K., Clare, E., Jones, G., Willerslev, E., Gilbert, M.T.P. 2011. Plos ONE 6(6): e21441. doi:10.1371/journal.pone.0021441.


Dietary preferences of the multimammate mouse (Mastomys natalensis, Smith 1832) across different habitats and seasons in Tanzania and Swaziland.

Mulungu, L.S., Massawe, A.W., Kennis, J., Crauwels, D., Eiseb, S., Mahlaba, T.A., Monadjem, A., Makundi, R.H., Katakweba, A.A.S., Leirs, H. & Belmain, S.R. 2011. Wildlife Research 38: 640-646.


Spatial and temporal population dynamics of rodents in three geographically different regions in Africa: implications for ecologically-based rodent management.

Massawe, A.W., Mulungu, L.S., Makundi, R.H., Dlamini, N., Eiseb, S.J., Kirsten, F., Mahlaba, T., Malebane, P., Von Maltitz, E., Monadjem, A., Taylor, P., Tutjavi, V. & Belmain, S.R. 2011. Africa Zoology 46: 393- 405.


Breeding season of Epomophorus walhbergi in the lowveld of Swaziland.

Monadjem, A. & Reside, A.E. Submitted. African Zoology


Roost use by two sympatric species of Scotophilus in a natural environment.

South African Journal of Wildlife Research

Monadjem, A., Raabe, T., Dickerson, B., Silvy, N. & McCleery, R. 2010.


A recent inventory of the bats of Mozambique with documentation of seven new species to the country

Acta Chiropterologica

Monadjem, A., Schoeman, M.C., Reside, A., Pio, D.V., Stoffberg, S., Bayliss, J., Cotterill, F.P.D., Curran, M., Kopp, M. & Taylor, P.J. 2010


Influence of rainfall on timing and success of reproduction in Marabou Storks


Monadjem, A. & Bamford, A.J.  (01-01-09)


Nesting habitat preference of the African white-backed vulture, Gyps africanus: a statistical model.


Bamford, A.J., Monadjem, A. & Hardy, I.W.  (01-01-09)


The diet of the aardwolf, Proteles cristatus at Malolotja Nature Reserve, western Swaziland

African Journal of Ecology

Matsebula, S.N., Monadjem, A., Roques, K.G. & Garcelon, D.K.  (01-01-09)


Development of non-explosive based methods for mass capture of vultures

South African Journal of Wildlife Research

Bamford, A.J., Diekmann, M., Monadjem, A. & Hardy, I.C.W.  (01-01-09)


Marabou Mania


Ara Monadjem, Brids of prey working group  (10-10-08)


Nest distribution and conservation status of eagles, selected hawks and owls in Swaziland


Monadjem, A. & Rasmussen, M.  (01-01-08)


The influence of riparian vegetation on the distribution and abundance of bats in an African savanna

Acta Chiropterologica

Monadjem, A. & Reside, A.  (01-01-08)


Dispersal of juvenile Marabou Storks Leptoptilos crumeniferus as determined by resightings


Monadjem, A., Bamford, A.J. & Rasmussen, M.  (01-01-08)

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Bats recorded from Koegelbeen Cave and selected other sites in the Northern Cape, South Africa

African Bat Conservation News  18: 2-4

Monadjem, A., Higgins, N., Smith, T. & Herrmann, E.  (01-01-08)


Georgaphical Distribution: Zygaspis vandami arenicola

African Herp News  46: 24-25

Litscha, T., Koen, C. and Monadjem, A.  (01-01-08)


Echolocation calls of rhinolophid and hipposiderid bats in Swaziland

South African Journal of Wildlife Research  37: 9-15

Monadjem, A., Reside, A. & Lumsden, L.  (01-01-07)


Nest success and conservation status of the Blue Swallow Hirundo atrocaerulea in Swaziland

Bird Conservation International  16

Monadjem, A., Boycott, R.C., Roques, K., Gama,R. & Garcelon, D.  (01-01-06)


Aspects of the breeding biology of the Chinspot Batis Batis molitor in Acacia savanna in Swaziland

Ostrich  77

Monadjem, A.  (01-01-06)


Longevity and movement of the common slit-faced bat Nycteris thebaica

African Bat Conservation News  9: 7

Monadjem, A.  (01-01-06)


Survival and roost-site selection in the African bat Nycteris thebaica (Chiroptera: Nycteridae) in Swaziland

Belgian Journal of Zoology  135: 103-107

Monadjem, A.  (01-01-05)


A Field Assessment of Priority Protection Worthy Areas of Swaziland: Makhonjwa, Manzimnyame, Sibebe and Nyonyane.

Special report to the Government of Swaziland.

Roques, K.G., Dobson, L., Dlamini, S.D., Monadjem, A., Boycott, R., Mahlaba, T. and Raw, D.  (19-06-03)


A Rapid Field Assessment of Protection Worthy Areas of Swaziland

Special report to the Government of Swaziland.

Roques, K.G.  (19-06-02)


Recent Publications


Taylor, P.J. Downs, S., Monadjem, A., Eiseb, S.J. Mulungu, L.S., Massawe, A.W., Mahlaba, T.A., Kirstin, F., Von Maltitz, E., Malebane, P., Makundi, R.H., Lamb, J. & Belmain, S.R. “Experimental treatment-control studies of ecologically based rodent management in Africa: balancing conservation and pest management.”, Wildlife Research 39: 51-61, 2012.


Lehmkuhl Noer, C., Dabelsteen, T., Bohmann, K. & Monadjem, A.

“Molossid bats in an African agro-ecosystem select sugar cane fields as foraging habitat.”, African Zoology 47: 1-11, 2012.


Morgan, T.C., McCleery, R.A., Moulton, M.P. & Monadjem, A.

“Are Southern Black Flycatchers (Melaenornis pammelaina) associated
with Fork-tailed Drongos (Dicrurus adsimilis)?”,
Ostrich 83: 109-111, 2012

Monadjem, A., Botha, A. & Murn, C.  “Survival of the African white-backed vulture Gyps africanus in north-eastern South Africa.”,African Journal of Ecology 51: 87-93, 2012.


Taylor, P.J., Stoffberg, S., Monadjem, A., Schoeman, M.C., Bayliss, J. & Cotterill, F.P.D.       “Four new bat species (Rhinolophus hildebrandtii complex) reflect Plio-Pleistocene divergence ofdwarfs and giants across an Afromontane Archipelago.”,  Plos ONE. 7(9): e41744. doi:10.1371/journal.pone.0041744, 2012.


Monadjem, A., Kane, A., Botha, A., Dalton, D. & Kotze, A. “Survival and     population dynamics of the marabou stork in an isolated population, Swaziland.” Plos ONE. 7(9): e46434.
doi:10.1371/journal.pone.0046434, 2012.


Monadjem, A., Virani, M.R., Jackson, C. & Reside, A. “Rapid decline and shift in the future distribution predicted for the endangered Sokoke Scops Owl Otus ireneae due to climate change.”, Bird Conservation International DOI: 10.1017/S0959270912000330, 2012.


Monadjem, A., Richards, L., Taylor, P.J. & Stoffberg, S. “High diversity of pipistrelloid bats (Vespertilionidae: Hypsugo, Neoromica and Pipistrellus) in a West African rainforest with the
description of a new species.”,
  Zoological Journal of the Linnean Society 167: 191-207, 2013.


Mdangia, M., Mulungu, L.S., Massawe, A.W., Eiseb, S., Tutjavic, V., Kirsten, F., Mahlaba, T., Malebane, P., Van Maltitz, E.,Monadjem, A., Dlamini, N., Makundi, R.H. & Belmain, S.R.  “Assessment of rodent damage to stored maize (Zea mays L.) on smallholder farms in Tanzania.”, International Journal of Pest Management DOI:10.1080/09670874.2012.744495, 2013.



Monadjem, A. & Reside, A.E.  “Breeding season of Epomophorus walhbergi in the lowveld of Swaziland.”, African Zoology 47: 321-325, 2013.


Long, A., Bailey, K., Greene, D.U., Tye, C., Parr, C., Lepage, H.K., Gielow, K.H., Monadjem, A. & McCleery, R.A. “Multi-scale habitat selection of the African Pygmy Mouse (Mus minutoides).”, African Journal of Ecology (early view), 2012.


Taylor, P.J., Monadjem, A., & Steyn, J. “Seasonal patterns of habitat use by insectivorous bats in a subtropical African agro-ecosystem dominated by macadamia orchards.”, African Journal of Ecology (early view), 2013.

Schoeman, M.C., Cotterill, F.P.D. (Woody), Taylor, P., & Monadjem, A. “Using potential distributions to explore environmental correlates of bat species richness in southern Africa: effects of model selection and taxonomy.”, Current Zoology , In Press.

Monadjem, A., McCleery, R.A. & Collier, B.A. “Activity and movement patterns of the tortoise Stigmochelys pardalis in a subtropical savanna.”, Journal of Herpetology, In press.

Denys, C., Kadjo, B., Missoup, A.D., Monadjem, A., & Aniskine, V. “New records of bats (Chiroptera) and karyotypes fromGuinean Mount Nimba (West Africa).”, Italian Journal of Zoology, In press.

Taylor, P.J., Sowler, S., Schoeman, M.C., & Monadjem, A. “Diversity of bats in the Soutpansberg and Blouberg Mountains of northern South Africa: complementarity of acoustic and non-acoustic survey methods.”, South African Journal of Wildlife Research, In


Book: Monadjem, A., Taylor, P., Cotterill, F.P.D. & Schoeman, M.C. 2010. Bats of
Southern and Central Africa: A Biogeographic and Taxonomic Synthesis.
University of the Witwatersrand, Johannesburg.




Marine Publications:

Developing a Code of Conduct for whale shark interactions in Mozambique.

Aquatic Conservation

Pierce, S.M., Mendez-Jimenez, A., Collins, K., Rosero, M. & Monadjem, A. 2010.


How large is the world’s largest fish? Measuring whale sharks Rhincodon typus with laser photogrammetry

Marine Ecology Progress Series

Rohner,C.A., Richardson, A.J., Marshall, A.D., Weeks, S.J. and Pierce, S.J. (Submitted)


Is whale shark tourism ecologically sustainable in Mozambique?

Marine Ecology Progress Series

Haskell, P.J., McGowan, A., Westling, A., Méndez-Jiménez, A., Rohner,C.A., Collins, K., Rosero-Caicedo, M., Salmond, J., Monadjem, A., Marshall, A.D., and Pierce, S.J. (Submitted)